Stevan J. Arnold ---- Publications


1. Arnold, S.J. 1972. Species densities of predators and their prey. American Naturalist 106: 220-236. pdf

2. Arnold, S.J. 1976. Sexual behavior, sexual interference and sexual defense in the salamanders Ambystoma maculatum, Ambystoma tigrinum, and Plethodon jordani. Zeitschrift für Tierpsychologie. 42: 247-300. pdf

3. Arnold, S.J. 1977. Polymorphism and geographic variation in the feeding behavior of the garter snake, Thamnophis elegans. Science 197: 676-678. pdf

4. Arnold, S.J. 1977. The courtship behavior of North American salamanders with some comments on Old World salamandrids. Pp. 141-183 IN: D. Taylor and S. Guttman (eds.), The Reproductive Biology of Amphibians. Plenum Press, New York. pdf

5. Arnold, S.J. 1978. Some effects of early experience on feeding responses in the common garter snake, Thamnophis sirtalis. Animal Behaviour. 26: 455-462. pdf

6. Arnold, S.J. 1978. The evolution of a special class of modifiable behaviors in relation to environmental pattern. American Naturalist. 112: 415-427. pdf

7. Arnold, S.J. and R.J. Wassersug. 1978. Differential predation on metamorphic anurans by garter snakes (Thamnophis): social behavior as a possible defense. Ecology 59: 1014-1022. pdf

8. Wade, M.J. and S. J. Arnold. 1980. The intensity of sexual selection in relation to male sexual behaviour, female choice and sperm precedence. Animal Behaviour 28: 446-461. pdf

9. Arnold, S.J. 1980. Inheritance and evolution of reproductive traits in garter snakes. American Zoologist. 20: 271 (Abstract). pdf

10. Arnold, S.J. 1981. The microevolution of feeding behavior. Pp. 409-453 IN: A. Kamil and T. Sargent (eds.), Foraging Behavior: Ecological, Ethological and Psychological Approaches. Garland Press, New York. pdf

11. Arnold, S.J. 1981. Behavioral variation in natural populations. I. Phenotypic, genetic and environmental correlations between chemoreceptive responses to prey in the garter snake, Thamnophis elegans. Evolution 35: 489-509. pdf   litter means

12. Arnold, S.J. 1981. Behavioral variation in natural populations. II. The inheritance of feeding response in crosses between geographic races of the garter snake, Thamnophis elegans. Evolution 35: 510-515. pdf

13. Arnold, S.J. 1981. Sociobiology evolving. Evolution 35: 824-825. (Book Review).

14. Arnold, S.J. 1982. A quantitative approach to antipredator performance: salamander defense against snake attack. Copeia 1982 (2): 247-253. pdf

15. Marx, H., G. Rabb and S.J. Arnold. 1982. Pythonodipsas and Spalerosophis: colubrid snake genera convergent to the vipers. Copeia 1982 (3): 553-561. pdf

16. Harvey, P.H. and S.J. Arnold. 1982. Female mate choice and runaway sexual selection. Nature 297: 533-534. pdf

17. Feder, M.E., and S.J. Arnold. 1982. Anaerobic metabolism and behavior during predatory encounters between snakes (Thamnophis elegans) and salamanders (Plethodon jordani). Oecologia 53: 93-97. pdf

18. Arnold, S.J. and L.D. Houck, 1982. Courtship pheromones: evolution by natural and sexual selection. pp. 173-211 IN: M. Nitecke (ed.), Biochemical Aspects of Evolutionary Biology. Univ. Chicago Press, Chicago. pdf

19. Kephart, D.G. and S.J. Arnold. 1982. Garter snake diets in a fluctuating environment: a seven year study. Ecology 63: 1232-1236. pdf

20. Arnold, S.J. 1983. Sexual selection: the interface of theory and empiricism. Pp. 67-107 IN: Mate Choice, P.P.G. Bateson (ed.), Cambridge Univ. Press. pdf

21. Arnold, S.J. 1983. Morphology, performance and fitness. American Zoologist 23: 347-361. pdf

22. Garland, T., Jr. and S.J. Arnold. 1983. The effect of a full stomach on locomotory performance of juvenile garter snakes (Thamnophis elegans). Copeia 1983: 1092-1096. pdf

23. Lande, R. and S.J. Arnold. 1983. The measurement of selection on correlated characters. Evolution 37: 1210-1226. pdf

24. Ayres, F.A. and S.J. Arnold. 1983. Behavioural variation in natural populations. IV. Mendelian models and heritability of a feeding response in the garter snake, Thamnophis elegans. Heredity 51: 405-413. pdf

25. Arnold, S.J. and A.F. Bennett. 1984. Behavioural variation in natural populations. III. Antipredator displays in the garter snake Thamnophis radix. Animal Behaviour 32:1108-1118. pdf

26. Arnold, S.J. and M.J. Wade. 1984. On the measurement of natural and sexual selection: theory. Evolution 38:709-719. pdf

27. Arnold, S.J. and M.J. Wade. 1984. On the measurement of natural and sexual selection: applications. Evolution 38:720-734. pdf

28. Houck, L.D., S.J. Arnold and R.A. Thisted. 1985. A statistical study of mate choice: sexual selection in a plethodontid salamander (Desmognathus ochrophaeus). Evolution 39: 370-386. pdf

29. Houck, L.D., S.G. Tilley and S.J. Arnold. 1985. Sperm competition in a plethodontid salamander: preliminary results. Journal of Herpetology 19: 420-423. pdf

30. Lande, R. and S.J. Arnold. 1985. Evolution of mating preference and sexual dimorphism. Journal of  Theoretical Biology 117: 651-664. pdf

31. Arnold, S.J. 1985. Quantitative genetic models of sexual selection. Experientia 41: 1296-1310. pdf

32. Arnold, S.J. 1986. Laboratory and field approaches to the study of adaptation. Pp. 157-179 IN: M. Feder and G. Lauder (eds.), Predator-prey Relationships. Univ. Chicago Press, Chicago. pdf

33. Arnold, S.J. 1986. Limits on stabilizing, disruptive and correlational selection set by the opportunity for selection. American Naturalist 128: 143-146. pdf

34. Arnold, S.J. 1986. Measuring selection. Science 232: 271-272 (Book Review). pdf

35. Arnold, S.J. and T. Halliday. 1986. Hyla regilla (Pacific Treefrog). Predation. Herpetological Review 17:44. pdf

36. Peterson, C.R. and S.J. Arnold. 1986. Individual variation in the thermoregulatory behavior of free-ranging garter snakes, Thamnophis elegans. American Zoologist 26:112A (Abstract).

37. Arnold, S.J. 1987. Natural selection in the wild. Animal Behaviour 35:308-309 (Book Review).

38. Arnold, S.J. 1987. The comparative ethology of courtship in salamandrid salamanders. 1. Salamandra and Chioglossa. Ethology 74:133-145. pdf

39. Heisler, L., M. Andersson, S.J. Arnold, C.R. Boake, G. Borgia, G. Hausfater, M. Kirkpatrick, R. Lande, J. Maynard Smith, P. O'Donald, A.R. Thornhill and F. Weissing. 1987. The evolution of mating preferences and sexually selected traits. Pp. 96-118 IN: J.W. Bradbury and M.B. Andersson (eds.), Sexual Selection: Testing the Alternatives. Dahlem Konferenzen. John Wiley and Sons, Chichester. pdf

40. Halliday, T.R. and S.J. Arnold. 1987. Multiple mating by females: a perspective from quantitative genetics. Animal Behaviour 35:939-941. pdf

41. Arnold, S.J. 1987. Quantitative genetic models of sexual selection: a review. Pp. 283-315 IN: S. Stearns (ed.), The Evolution of Sex and Its Consequences. Birkhauser, Basel. pdf

42. Arnold, S.J. 1987. Genetic correlation and the evolution of physiology. Pp. 189-212 IN: M.E. Feder, A.L. Bennett, W.W. Burggren and R.B. Huey (eds.), New Directions in Ecological Physiology. Cambridge Univ. Press. pdf

43. Arnold, S.J. 1988. Behavior, energy and fitness. American Zoologist 28:815-827. pdf

44. Houck, L.D., S.J. Arnold and A. Hickman. 1988. Tests for sexual isolation in plethodontid salamanders (genus Desmognathus).  Journal of Herpetology 22:186-191. pdf

45. Arnold, S.J. 1988. Snakes: ecology and evolutionary biology. (Book Review) Herpetologica 44: 259-260. pdf

46. Arnold, S.J. 1988. Quantitative genetics and selection in natural populations: microevolution of vertebral numbers in the garter snake Thamnophis elegans. Pp. 619-636 IN: B.S. Weir, E.J. Eisen, M.M. Goodman, and G. Namkoong (eds.), Proceedings of the Second International Conference on Quantitative Genetics. Sinauer, Sunderland, MA pdf

47. Arnold, S.J. and A.F. Bennett. 1988. Behavioural variation in natural populations. V. Morphological correlates of locomotion in the garter snake Thamnophis radix. Biological Journal of the Linnean Society 34: 175-190. pdf

48. Price, T., M. Kirkpatrick and S.J. Arnold. 1988. Directional selection and the evolution of breeding date in birds. Science 240: 798-799. pdf

49. Arnold, S.J. and T.R. Halliday. 1988. Multiple mating: natural selection is not evolution. Animal Behaviour 36: 1547-1548. pdf

50. Lynch, M. and S.J. Arnold. 1988. The measurement of selection on size and growth. Pp. 47-59 IN: B. Ebenman and L. Persson (eds.), Size-structured Populations, Ecology and Evolution. Springer-Verlag, Berlin. pdf

51. Arnold, S.J. 1989. Biology of the Reptilia, vol. 16 (Book Review). American Scientist 77: 187.

52. Arnold, S.J., P. Alberch, V. Csányi, R.C. Dawkins, S.B. Emerson, B. Fritzsch, T.J. Horder, J. Maynard Smith, M.J. Starck, E.S. Vrba, G.P. Wagner and D.B. Wake. 1989. How do complex organisms evolve? Pp. 403-433 IN: D.B. Wake and G. Roth (eds.), Complex Organismal Functions: Integration and Evolution in Vertebrates. Wiley, New York. pdf

53. Emerson, S.B. and S.J. Arnold. 1989. Intra- and interspecific relationships between morphology, performance, and fitness. Pp. 295-314 IN: D.B. Wake and G. Roth (eds.), Complex Organismal Functions: Integration and Evolution in Vertebrates. Wiley, New York. pdf

54. Huey, R.B., C.R. Peterson, S.J. Arnold and W.P. Porter. 1989. Hot rocks and not-so-hot rocks: thermal consequences of retreat site selection by garter snakes. Ecology 70: 931-944. pdf

55. Phillips, P.C. and S.J. Arnold. 1989. Visualizing multivariate selection. Evolution 43: 1209-1222. pdf

56. Verrell, P.A. and S.J. Arnold. 1989. Behavioral observations on sexual isolation between allopatric populations of the Mountain Dusky Salamander, Desmognathus ochrophaeus. Evolution 43: 745-755. pdf

57. Arnold, S.J. and C.R. Peterson. 1989. A test for temperature effects on the ontogeny of shape in the garter snake Thamnophis sirtalis. Physiological Zoology 62: 1316-1333. pdf

58. Arnold. S.J. 1990. Inheritance and the evolution of behavioral ontogenies. Pp. 167-189 IN: M. Hahn, J. Hewitt, N. Henderson, and R. Benno (eds.), Developmental Behavior Genetics: Neural, Biometrical, and Evolutionary Approaches. Oxford Univ. Press, New York. pdf

59. Kirkpatrick, M., T. Price and S.J. Arnold. 1990. The Darwin-Fisher theory of sexual selection in monogamous birds. Evolution 44:180-193. pdf

60. Tilley, S.G., P.A. Verrell and S.J. Arnold. 1990. Correspondence between sexual isolation and allozyme differentiation: a test in the salamander Desmognathus ochrophaeus. Proceedings of the National Academy of Sciences U.S.A. 87:2715-2719. pdf

61. Arnold, S.J. 1990. Reproductive success: studies of individual variation in contrasting breeding systems. (Book Review). American Scientist 78:468.

62. Arnold, S.J. and T. Halliday. 1991. Multiple mating by females: design and interpretation of selection experiments. Animal Behaviour 43:178-179. pdf

63. Duvall, D., S.J. Arnold and G.W. Schuett. 1992. Pitviper mating systems: ecological potential, sexual selection, and microevolution. Pp. 321-336 IN: J.A. Campbell & E.D. Brodie, Jr. (eds.), The Biology of the Pitvipers. Selva Press, Tyler, TX. pdf

64. Arnold, S.J. 1992. Behavioural variation in natural populations. VI. Prey responses by two species of garter snakes in three regions of sympatry. Animal Behaviour 44:705-719. pdf

65. Arnold, S.J. 1992. Constraints on phenotypic evolution. American Naturalist 140:S85-S107. pdf

66. Arnold, S.J. 1993. Foraging theory and prey size - predator size relations in snakes. Pp. 87-115 IN: R.A. Seigel & J.T. Collins (eds.), Snakes: Ecology and Behavior. McGraw Hill, New York. pdf

67. Duvall, D., G.W. Schuett and S.J. Arnold. 1993. Ecology and evolution of snake mating systems. Pp. 165-200 IN: R.A. Seigel & J.T. Collins (eds.), Snakes: Ecology and Behavior. McGraw Hill, New York. pdf

68. Arnold, S. J., N. L. Reagan and P. A. Verrell. 1993. Reproductive isolation and speciation in plethodontid salamanders. Herpetologica 49: 216-228. pdf

69. Arnold, S.J. and D. Duvall. 1994. Animal mating systems: a synthesis based on selection theory. American Naturalist 143:317-348. pdf

70. Arnold, S. J. 1994. Is there a unifying concept of sexual selection that applies to both plants and animals? American Naturalist 144:S1-S12. pdf

71. Arnold, S. J. 1994. Bateman's principles and the measurement of sexual selection in plants and animals. American Naturalist 144:S126-S149. pdf

72. Arnold, S.J. 1994. Multivariate inheritance and evolution: a review of concepts. Pp. 17-48 IN: C.R.P. Boake (ed.), Quantitative Genetics Studies of the Evolution of Behavior. Univ. Chicago Press, Chicago. pdf

73. Arnold, S.J. 1994. Constraints on phenotypic evolution. pp. 258-278 IN: L.A. Real (ed.), Behavioral Mechanisms in Evolutionary Biology. Univ. Chicago Press, Chicago. pdf

74. Arnold, S.J. 1995. Monitoring quantitative genetic variation and evolution in captive populations. pp. 295-317 IN: J. Ballou, M. Gilpin & T. Foose (eds.), Population Management for Survival and Recovery: Analytical Methods and Strategies in Small Population Conservation. Columbia University Press. pdf

75. Arnold, S. J., C. R. Peterson and J. Gladstone. 1995. Behavioural variation in natural populations. VII. Maternal body temperature does not affect juvenile thermoregulation in a garter snake (Thamnophis elegans). Animal Behaviour 50: 623-633. pdf

76. Arnold, S. J. 1995. Fauna of Australia. Vol. 2A, Amphibia and Reptilia. (Book Review) Copeia 1995: 247-248. pdf

77. Webster, M. S., S. Pruett-Jones, D. F. Westneat, and S. J. Arnold 1995. Measuring the effects of pairing success, extra-pair copulations and mate quality on the opportunity for sexual selection. Evolution 49: 1147-1157. pdf

78. Arnold, S. J., P. A. Verrell and S. G. Tilley. 1996. The evolution of asymmetric sexual isolation: a polygenic model and a test case. Evolution 50:1024-1033. pdf   Excel file that calculates standard errors of isolation measures    Readme file with instructions

79. Arnold, S. J. and H. J. Brockmann. 1996. Evolution of behavior, approaches to studying behavioral change. Pp. 673-682 IN: L. D. Houck & L. C. Drickamer (eds.), Foundations of Animal Behavior. Univ. Chicago Press, Chicago. pdf

80. Kelley, K. C., S. J. Arnold and J. Gladstone. 1997. The effects of substrate and vertebral number on locomotion in the garter snake Thamnophis elegans. Functional Ecology 11:189-198. pdf

81. Arnold, S. J. 1998. Snakes, the evolution of mystery in nature. (Book Review). Integrative and Comparative Biology 1:76-77.

82. Bronikowski, A. M. and S. J. Arnold. 1999. The evolutionary ecology of life history variation in the garter snake Thamnophis elegans. Ecology 80:2314-2325. pdf

83. Bernardo, J. and S. J. Arnold. 1999. Mass-rearing of plethodontid salamander eggs. Amphibia-Reptilia 20:219-224.pdf

84. Phillips, P. C. and S. J. Arnold. 1999. Hierarchial comparison of genetic variance-covariance matrices. I. Using the Flury hierarchy. Evolution 53:1506-1515. pdf   Software

85. Arnold, S. J. and P. C. Phillips. 1999. Hierarchial comparison of genetic variance-covariance matrices.II. Coastal-inland divergence in the garter snake, Thamnophis elegans. Evolution 53:1516-1527. pdf

86. Osypka, N. M. and S. J. Arnold. 2000. The developmental effect of sex ratio on a sexually dimorphic scale count in the garter snake Thamnophis elegans. Journal of Herpetology 34:1-5. pdf

87. Arnold, S. J. 2000. Systematics at the turn of a century. Pp. 167-178 IN: R. C. Bruce, R. G. Jaeger, and L. D. Houck (eds.), The Biology of Plethodontid Salamanders. Kluwer/Plenum, New York, New York. pdf

88. Jones, A. G., G. Rosenqvist,  A. Berglund,  S. J. Arnold and J. C. Avise. 2000. The Bateman Gradient and the cause of sexual selection in a sex-role-reversed pipefish. Proceedings of the Royal Society of London B: Biological Sciences 267:1-4. pdf

89. Arnold, S. J., J. Kagan and B. Taylor. 2000. Summary of current status of Oregon’s biodiversity. Pp.121-162 IN: The Oregon State of the Environment Report 2000. pdf

90. Arnold, S. J. and J. A. Anthony. 2000. Summary of the current status of exotic species in Oregon. Pp. 127-131 IN: Oregon State of the Environment Report 2000. pdf

 91. Bronikowski, A. M. and S. J. Arnold. 2001. Cytochrome b phylogeny does not match subspecific classification in the western terrestrial garter snake, Thamnophis elegans. Copeia 2001: 508-513. pdf

92. Alfaro, M. and S. J. Arnold. 2001. Molecular systematics and evolution of Regina and the thamnophiine snakes. Molecular Phylogenetics and Evolution 21: 408-423. pdf

93. Arnold, S. J., M. E. Pfrender, A. G. Jones. 2001. The adaptive landscape as a conceptual bridge between micro- and macroevolution. Genetica 112/113: 9-32. pdf

94. Jones, A. G., M. S. Blouin and S. J. Arnold. 2001. Genetic variation in two populations of the rough-skinned newt (Taricha granulosa) assessed using novel tetranucleotide microsatellite loci. Molecular Ecology Notes 1: 293-296. pdf

95. Arnold, S. J. and C. R. Peterson. 2002. A model for optimal reaction norms: the case of the pregnant garter snake and her temperature sensitive embryos. American Naturalist 160: 306-316. pdf

96. Jones, A. G., E. M. Adams and S. J. Arnold. 2002. Topping off: a mechanism of sperm competition in a vertebrate. Proceedings of the National Academy of Sciences U.S.A. 99: 2078-2081. pdf

97. Boake, C. R. B., S. J. Arnold, F. Breden, L. Meffert, M. Ptacek, M. Ritchie, B. Taylor, J. B. Wolf, and A. J. Moore.  2002.  Genetic tools for studying adaptation and the evolution of behavior. American Naturalist 160(Suppl.): S143-S159. pdf

98. Jones, A. G., J. R. Arguello and S. J. Arnold. 2002.  Validation of Bateman’s principles: a genetic study of sexual selection and mating patterns in newts.  Proceedings of the Royal Society of London B: Biological Sciences 269: 2533-2539. pdf

99.  Arnold, S. J., M. E. Pfrender, A. G. Jones. 2002. The adaptive landscape as a conceptual bridge between micro- and macroevolution. Pp. 9-32 IN: A. P. Hendry and M. T. Kinnison (eds.). Microevolution – Rate, Pattern, Process. Kluwer Academic Publishers, Dordrecht. pdf

100.  Bernardo, J. and S. J. Arnold. 2002. Gyrinophilus porphyriticus (Spring Salamander). Male combat. Herpetological Review 33: 121-122. pdf

101.  Houck, L. D. and S. J. Arnold. 2003. Courtship and mating behavior. Pp. 383-424 IN:  D. Sever (ed.), Reproductive Biology and Phylogeny of Urodela.  M/s Science Publications, Endfield, NH. pdf

102.  Jones, A. G., S. J. Arnold and R. Bürger.  2003.  Stability of the G-matrix in a population experiencing mutation, stabilizing selection, and genetic drift.  Evolution 57: 1747-1760. pdf

103.  Arnold, S. J. 2003. Too much natural history, or too little?  Animal Behaviour 65: 1065-1068. pdf

104.  Arnold, S. J. 2003.  Performance surfaces and adaptive landscapes. Integrative & Compartive Biology 43: 367-375. pdf

105.  Jones, A. G., J. R. Arguello, and S. J. Arnold. 2004.  Molecular parentage analysis in experimental newt populations:  the response of mating system measures to variation in the operational sex ratio.  American Naturalist 164: 444-456. pdf

106.  Mead, L. S. and S. J. Arnold.  2004.  Quantitative genetic models of sexual selection.  Trends in Ecology and Evolution 19: 264-271. pdf

 107.  Watts R., C. Palmer, R. Feldhoff, P. Feldhoff, L. D. Houck, A. Jones, M. Pfrender, S. Rollmann, and S. J. Arnold.  2004.  Stabilizing selection on behavior and morphology masks positive selection on the signal in a salamander pheromone signaling complex.  Molecular Biology & Evolution 21: 1032-1041. pdf

 108.  Jones, A. G., S. J. Arnold, and R. Bürger. 2004. Evolution and stability of the G-matrix on a landscape with a moving optimum.  Evolution 58: 1639-1654. pdf

109.  DeGross, D. J., L. S. Mead, and S. J. Arnold. 2004.  Novel tetranucleotide microsatellite markers from the Del Norte Salamander (Plethodon elongatus) with applications to its sister species the Siskiyou Mtn. Salamander (P. stormi).  Molecular Ecology Notes 4: 352-354. pdf

110.   Adams, E. M., A. G. Jones, and S. J. Arnold. 2005.  Multiple paternity in a natural population of a salamander with long-term sperm storage.  Molecular Ecology 14: 1803-1810. pdf

111.   Palmer, C., R. A. Watts, R. Gregg, M. McCall, L. D. Houck, R. Highton, and S. J. Arnold. 2005.  Lineage-specific differences in evolutionary mode in a salamander courtship pheromone.  Molecular Biology & Evolution 22: 2243-2256. pdf

112.   Manier, M. K. and S. J. Arnold.  2005.  Population genetic analysis identifies source-sink dynamics for two sympatric garter snake species (Thamnophis elegans and T. sirtalis).  Molecular Ecology 14: 3965-3976. pdf

113.  O’Donnell, R. P. and S. J. Arnold.  2005.  Evidence for selection on thermoregulation:  effects of temperature on duration of pregnancy and embryo mortality in the garter snake Thamnophis elegans. Copeia 2005: 929-933. pdf

114. Arnold, S. J. 2005.  The utlimate causes of phenotypic integration: lost in translation. (Book Review). Evolution 59: 2059-2061. pdf

115. Manier, M. K. and S. J. Arnold. 2006.  Ecological correlates of population genetic structure: a comparative approach using a vertebrate metacommunity.  Proceedings of the Royal Society of London B: Biological Sciences 273: 3001-3009. pdf

116.  Estes, S. and S. J. Arnold. 2007.  Resolving the paradox of stasis: models with stabilizing selection explain evolutionary divergence on all timescales.  American Naturalist 169: 227-244. pdf supplementary material.

117.  Palmer, C. A., R. A Watts, L. D. Houck, A. Picard and S. J. Arnold. 2007.  Evolutionary replacement of components in a salamander pheromone signaling complex: more evidence for phenotypic-molecular decoupling. Evolution 61: 202-215. pdf

118.  Houck, L. D.,  C. A. Palmer, R. A. Watts, S. J. Arnold, P. W. Feldhoff and R. C. Feldhoff.  2007.  A new vertebrate courtship pheromone, PMF, affects female receptivity in a terrestrial salamander. Animal Behaviour 73: 315-320. pdf

119. Houck, L. D., L. S. Mead, R. A. Watts, S. J. Arnold, P. W. Feldhoff and R. C. Feldhoff.  2007.  A candidate vertebrate pheromone, SPF, increases female receptivity in a salamander.  Pp. 213-221 IN:  J. Hurst, R. Beynon, and D. Muller-Schwarze (eds.), Chemical Signals in Vertebrates  11. pdf

120.  Sparkman, A. M., S. J Arnold and A. M. Bronikowski.  2007.  An empirical test of evolutionary theories for reproductive sensescence and reproductive effort in the garter snake Thamnophis elegansProceedings of the Royal Society of London B: Biological Sciences 274: 943-950. pdf

121. Manier, M. K., C. M. Seyler, and S. J. Arnold. 2007.  Adaptive divergence within and between ecotypes of the terrestrial garter snake, Thamnophis elegans, assessed with Fst-Qst comparisons.  Journal of Evolutionary Biology 20: 1705-1719. pdf  Figure 1a  Figure 1b Appendix S1 Appendix S2

122.  Jones, A. G., S. J. Arnold and R. Bürger. 2007.  The mutation matrix and the evolution of evolvability.  Evolution 61: 727-745. pdf

123.  Palmer, C. A., D. M. Hollis, R. A. Watts, L. D. Houck, M. A. McCall, R. G. Gregg and S. J. Arnold.  2007.  Plethodontid modulating factor (PMF), a hypervariable salamander courtship pheromone in the three-finger protein superfamily.  FEBS Journal 274: 2300-2310. pdf

124.  Hohenlohe, P. A. and S. J. Arnold.  2008.  MIPoD: a hypothesis-testing framework for microevolutionary inference from patterns of divergence.  American Naturalist 171: 366-385. pdf Appendix MIPoD software

125.  Houck, L. D., R. A. Watts, S. J. Arnold, K. E. Bowen, K. M. Kiemnec, H. A. Godwin, P. W. Feldhoff and Richard C. Feldhoff.  2008.  A recombinant courtship pheromone affects sexual receptivity in a plethodontid salamander.  Chemical Senses 33: 623-631. pdf

126.  Stinchcombe, J. R., A. F. Agrawal, P. A. Hohenlohe, S. J. Arnold, and M. W. Blows.  2008.  Estimating nonlinear selection gradients using quadratic regression coefficients: double or nothing?  Evolution 62: 2435-2440. pdf

127.  Arnold, S. J., R. Bürger, P. A. Hohenlohe, B. C. Ajie and A. G. Jones.  2008.   Understanding the evolution and stability of the G-matrix.  Evolution 62: 2451-2461. pdf spreadsheet

128. Kiemnec-Tyburczy, K. M., R. A. Watts, R. G. Gregg, D. von Borstel, and S. J. Arnold.  2009.  Evolutionary shifts in courtship pheromone composition revealed by EST analysis of plethodontid salamander mental glands. Gene 432:75-81. pdf

129. Uyeda, J. C., S. J. Arnold, P. A. Hohenlohe, and L. S. Mead.  2009.  Drift promotes speciation by sexual selection.  Evolution 63: 583-594. pdf  Explications and simulation run examples Supplementary appendix and tables Errata

130. Arnold, S. J. 2009.  Russell Lande. Pp. 675-676, IN: M. J. Ruse and J. Travis (eds.), Evolution, the First Four Billion Years, Harvard University Press, Cambridge, MA. pdf 

131.  Palmer, C.A., A. Picard, R. A. Watts, L. D. Houck and S. J. Arnold. 2010. Rapid evolution of Plethodontid Modulating Factor (PMF), a hypervariable salamander courtship pheromone, is driven by positive selection. Journal of Molecular Evolution 70: 427-440. pdf 

 132. Barker, B. S., S. J. Arnold, and P. C. Phillips. 2010.  A test of the conjecture that G-matrices are more stable than B-matrices.  Evolution 64: 2601-2613. pdf

 133. Hohenlohe, P. A. and S. J. Arnold. 2010.  The dimensionality of mate choice, sexual isolation and speciation.  Proceedings of the National Academy of Sciences U.S.A. 107: 16583-165588. pdf  Supplementary information

 134. Kiemnec-Tyburczy, K. M., R. A. Watts, and S. J. Arnold. 2011.  Characterization of two putative cytokine receptors, gp130 and ciliary neutrophic factor receptor, from terrestrial salamanders.  Genes & Genetic Systems 86: 131-137. pdf

 135. Uyeda, J. C., T. F. Hansen, S. J. Arnold, and J. Pienaar.  2011. The million-year wait for macroevolutionary bursts.  Proceedings of the National Academy of Sciences U.S.A. 108: 15908-15913. pdf   Supplementary information

 136. Miller, D. A., W. R. Clark, S. J. Arnold, and A. M. Bronikowski.  2011Stochastic population dynamics and life-history evolution in the western terrestrial garter snake. Ecology 92: 1658-1671. pdf

 137. Kiemnec-Tyburczy, K. M., S. K. Woodley, R. A. Watts, S. J. Arnold, and L. D. Houck. 2011. Expression of vomeronasal receptors and related signaling molecules in the nasal cavity of a caudate  amphibian (Plethodon shermani).  Chemical Senses doi: 10.1093/chemse/bjr105. pdf 

138. Highton, R., A. P. Hastings, C. A. Palmer, R. A. Watts, C. A. Hass, M. Culver, and S. J. Arnold. 2012.  Concurrent speciation in the Eastern Woodland Salamanders (genus Plethodon): DNA sequences of the complete albumin and partial 12s mitochondrial genes.  Molecular Phylogenetics & Evolution 63: 278-290. pdf 

139.  Jones, A. G., S. J. Arnold, R. Bürger, P. A. Hohenlohe, and J. C. Uyeda. 2012.  The effects of stochastic and episodic movement of the optimum on the evolution of the G-matrix and the response of the trait mean to selection.  Journal of  Evolutionary Biology 25: 2210-2231. pdf

140.  Losos, J. B., S. J. Arnold, G. Bejerano, E.D. Brodie III, D. Hibbett, H. E. Hoekstra, D. P. Mindell, A. Monteiro, C. Moritz, H. A. Orr, D. A. Petrov, S. S. Renner, R. E. Ricklefs, P. S. Soltis, and T. L. Turner.  2013.  Evolutionary biology for the 21st century. PLOS Biology 11(1): e1001466. doi:10.1371/journal.pbio.1001466. pdf   Supporting Information 

141.  Sparkman, A. M., J. G. Billings, D. von Borstel, A. M. Bronikowski, and S. J. Arnold.  2013.  Avian predation and the evolution of life-histories in the garter snake Thamnophis elegans.  American Midland Naturalist 170: 66-85. pdf Supplementary Information

142. Friesen, C. R., R. T. Mason, S. J. Arnold, and S. R. Estes. 2013.  Patterns of sperm use in two populations of Red-sided Garter Snake (Thamnophis sirtalis parietalis) with long-term female sperm storage.  Canadian Journal of Zoology 92: 33-40. pdf

143. Arnold, S. J. 2014. Phenotypic evolution: the ongoing synthesis.  The American Naturalist 183:729-746. pdf Appendix

144.  Jones, A. G., R. Bürger, and S. J. Arnold. 2014.  Epistasis and natural selection shape the mutational architecture of complex traits.  Nature Communications DOI: 10.1038/ncomms4709. pdf  Supplementary Information

145.  Wilburn, D. B., S. L. Eddy, A. J. Chouinard, S. J. Arnold, R. C. Feldhoff, and L. D. Houck. 2015. Pheromone isoform composition differentially affects female behaviour in the red-legged salamander, Plethodon shermani.  Animal Behaviour 100:1-7. pdf

146.  Arnold, S. J. and L. D. Houck.  2016.  Can the Fisher-Lande process account for birds-of-paradise and other sexual radiations?  The American Naturalist 187:717-735. pdf  Supplemental figures ppt

147.  Pierson, T. W., S. J. Arnold, M. K. Hamed, W. Lattea, and E. T. Carter. 2017.  Courtship behavior of the Yonahlossee Salamander, Plethodon yonahlossee: observations in the field and laboratory. Herpetological Conservation & Biology 12:1-15. pdf

148.  Doten, K., G. W. Bury, M. Rudenko, and S. J. Arnold.  2017.  Courtship in the Torrent Salamander, Rhyacotriton, has an ancient and stable history.  Herpetological Conservation & Biology 12:457-469. pdf  interview

149.  Arnold, S. J., K.  Kiemnec-Tyburczy, and L. D. Houck.  2017.  The evolution of courtship behavior in plethodontid salamanders, contrasting patterns of diversification and stasis.  Herpetologica 73:190-205. pdf  Appendix S1  Appendix S2

150.  Wilburn, D. B., L. D. Houck, S. J. Arnold, P. W. Feldhoff, and R. C. Feldhoff.  2017.  Gene duplication, co-option, structural evolution, and phenotypic tango in plethodontid courtship pheromones.  Herpetologica 73:206-219. pdf

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